diff --git a/.pql/changelog/ticket_history/2026-07.sql b/.pql/changelog/ticket_history/2026-07.sql
index 34ccb23d4..5895714ef 100644
--- a/.pql/changelog/ticket_history/2026-07.sql
+++ b/.pql/changelog/ticket_history/2026-07.sql
@@ -205,3 +205,86 @@ Chirality decisions resolved (2026-07-02, with Jeroen):
 NAMING CONFIRMED: Keid''s habitable body Cadwal → Linnaeus (the great classifier; ties to Solander/Solandar taxonomy + the ''five kingdoms''). Capital stays Vance.', NULL, '2026-07-02 21:24:12', '2026-07-02 21:24:12.734', '2026-07-02 21:24:12.734', NULL, 'b13c9f1ce1d47437fdb599e6efce7f0e', 2) ON CONFLICT(hash) DO NOTHING;
 INSERT INTO ticket_history (ticket_record_id, field, old_value, new_value, changed_by, changed_at, created_at, updated_at, deleted_at, hash, canonical_version) VALUES ('06FJ91R8RV38BJXGB2V55FP3B8', 'status', 'backlog', 'in_progress', NULL, '2026-07-03 20:18:52', '2026-07-03 20:18:52.241', '2026-07-03 20:18:52.241', NULL, '89aa6692b3a1ab821cb7b02c3970f5a5', 2) ON CONFLICT(hash) DO NOTHING;
 INSERT INTO ticket_history (ticket_record_id, field, old_value, new_value, changed_by, changed_at, created_at, updated_at, deleted_at, hash, canonical_version) VALUES ('06FJ91R8RV38BJXGB2V55FP3B8', 'status', 'in_progress', 'in_progress', NULL, '2026-07-03 20:20:01', '2026-07-03 20:20:01.466', '2026-07-03 20:20:01.466', NULL, '6d2a3d77d702940bbc9207e4b9ce2b26', 2) ON CONFLICT(hash) DO NOTHING;
+INSERT INTO ticket_history (ticket_record_id, field, old_value, new_value, changed_by, changed_at, created_at, updated_at, deleted_at, hash, canonical_version) VALUES ('06FJ91R8RV38BJXGB2V55FP3B8', 'description', 'Operationalize the D-247 native-biosphere premise. Full brief appended.
+
+---
+BRIEF (2026-07-02) — governing record: D-247 (governance/decisions/content.md); consumer: D-246 / T-1084 mosaic register.
+
+WHAT THIS IS
+D-247 established that habitable Reach worlds were alive before humans (the settlement timeline forbids terraforming from dead rock), and that an AUTHORED per-world chirality bit gates whether native life is edible. This ticket turns that canon into data + reconciled wiki + the vocabulary the mosaic needs. Owner: Miri (worldbuilding/continuity). Stakeholder: Burnelli (food-import economics). Co-maintainer for the vocab: Araminta (visual grammar, per the T-995 ObjectTag precedent).
+
+THE MODEL (locked in D-247 — do not re-litigate, apply it):
+- `biosphere_class ∈ {NativeCompatible, NativeMirror, TerraformedSterile, Airless}`. It is an AUTHORED INPUT, never seed-derived — it follows a world''s economic/cultural identity.
+- Compatible ("levo") = edible; coupling *permits* an authored hazard but does not force it (most are benign gardens).
+- Mirror ("dextro") = inedible but *guaranteed* inert (can''t infect/poison — safe to study).
+- "Safe" ≠ "edible" — decoupled. Bio-safe is guaranteed on Mirror; food-safe is Compatible-only.
+- Achiral seam (always works either way): breathe air, drink water, mine ore, burn/build with native biomass. Only eat/metabolize/medicate/infect/chirally-poison are gated.
+- ~50/50 Compatible:Mirror is a DESIGN TARGET across canon-silent worlds, not a per-world draw.
+
+WORKSTREAMS
+1. CHIRALITY ASSIGNMENT (Miri). Give every inhabited body a biosphere_class by authored identity: principled default from `economic_base_primary` + `planet_class` (food/drink exporters → Compatible; dead+maintained-atmosphere → TerraformedSterile), explicit overrides where canon speaks. Locked exemplars: Arbour (GJ-338B)=Compatible; Solandar (GJ-827)=Mirror; Keid (GJ-166A)=Compatible (preserved *from exploitation* of edible alien life — the stronger conservation motive; Solandar is the Mirror-inert preserve); Oshima (GJ-79)=TerraformedSterile. Hit the ~50/50 target across the silent remainder.
+2. WIKI CONTINUITY (Miri). Reconcile the entries that already gesture at native biochemistry so each is consistent with its assigned bit + the mechanism: GJ-827, GJ-524, GJ-166A, GJ-684A, GJ-197, GJ-79, GJ-124 (resolve whether Hạ Long''s "protein" is native-Compatible harvest or Earth aquaculture). Additive/explanatory, not contradictory — preserve deliberately-vague lines as intentional.
+3. biosphere_class AUTHORING PATH (asset-pipeline discipline — schema + source + values together, per .claude/rules/asset-pipeline.md). Add `biosphere_class` to wiki body frontmatter (where `substrate:` already lives), a nullable `bodies.biosphere_class` column in systems-schema.sql + MIGRATION_SQL, and importer read. NOT needed for D-246; lands with this pass.
+4. ALIEN-ECOLOGY VOCABULARY (Miri + Araminta) — feeds T-1084 directly. The mosaic palette is keyed on surface-class × climate-biome × biosphere-register (D-246 Palette-key). Provide the micro-habitat NAMES per biome × register: native-mirror niches get *alien* clade names (Solandar''s "five kingdoms" flavor); introduced-Earth niches get Earth names (wheatfield, orchard, pasture). Fill the fixed niche spine (canopy / understory / wet-hollow / pioneer / bare) per climate-biome (savanna gallery-copse, tundra thermokarst / lichen-heath, boreal muskeg / deadfall, …). DETERMINISM: the vocab must be fixed before T-1084 generates its first world (a later addition re-rolls affected voxels). Table size is not a perf/storage concern — author generously but to taste (~100k is headroom, not a target).
+
+BURNELLI (stakeholder, follow-on): a food-supply demand primitive keyed on biosphere_class (Mirror + frontier → import dependency; Compatible → local-food capable). Phase-2 economy consumer; does not block this ticket.
+
+CONSTRAINTS
+- Authored input, not derivation (Arbour''s estate food must never be coin-flipped inedible).
+- Semantic axes stay clade-agnostic (Forest = canopy whether oak or mega-fungus); the register skins niches, it does not change gameplay semantics.
+- The mosaic geometry is chirality-blind — this work does NOT touch voxel geometry, only edibility/economy/naming/vocab.
+
+Chirality decisions resolved (2026-07-02, with Jeroen):
+- Locked exemplars: Arbour=NativeCompatible, Solandar=NativeMirror, Keid=NativeCompatible, Oshima=TerraformedSterile.
+- Miri assignments accepted: GJ-524=Mirror, Freyburg(GJ-684A)=Compatible, Hạ Long(GJ-124)=Compatible (native-Compatible harvest, per her rationale).
+- USER OVERRIDES: Puerto Último(GJ-197)=NativeCompatible (breadbasket identity over Miri''s balance-motivated Mirror). Edict/Sirius(GJ-244A)=Airless — keep the world harsh/frozen (deliberately-inconvenient below-grade transit capital is load-bearing canon); no climate change; below-grade sealed habitat = no open-air biosphere.
+- Two-gate default rule ACCEPTED for canon-silent worlds: Gate 0 (habitability from atmosphere/hydrosphere frontmatter → Airless / TerraformedSterile / Alive), Gate 1 (Compatible vs Mirror from economic_base_primary; agriculture→Compatible near-lock, penal→Mirror, research→50/50 lever).
+- NAMING: Keid''s habitable body renamed Cadwal→(pending final pick; rec Linnaeus — pairs with Solander, its student naturalist; or Beagle). Capital stays ''Vance'' (already the author tribute). Miri''s IP flag (Cadwal = Vance''s exact book title) is the driver.
+- Still to APPLY: the 7 additive wiki continuity edits (Miri specified exact text), the biosphere_class frontmatter+schema+importer plumbing (workstream 3), and the Cadwal→<name> rename across wiki + the GJ166A atlas proposal.
+
+NAMING CONFIRMED: Keid''s habitable body Cadwal → Linnaeus (the great classifier; ties to Solander/Solandar taxonomy + the ''five kingdoms''). Capital stays Vance.', 'Operationalize the D-247 native-biosphere premise. Full brief appended.
+
+---
+BRIEF (2026-07-02) — governing record: D-247 (governance/decisions/content.md); consumer: D-246 / T-1084 mosaic register.
+
+WHAT THIS IS
+D-247 established that habitable Reach worlds were alive before humans (the settlement timeline forbids terraforming from dead rock), and that an AUTHORED per-world chirality bit gates whether native life is edible. This ticket turns that canon into data + reconciled wiki + the vocabulary the mosaic needs. Owner: Miri (worldbuilding/continuity). Stakeholder: Burnelli (food-import economics). Co-maintainer for the vocab: Araminta (visual grammar, per the T-995 ObjectTag precedent).
+
+THE MODEL (locked in D-247 — do not re-litigate, apply it):
+- `biosphere_class ∈ {NativeCompatible, NativeMirror, TerraformedSterile, Airless}`. It is an AUTHORED INPUT, never seed-derived — it follows a world''s economic/cultural identity.
+- Compatible ("levo") = edible; coupling *permits* an authored hazard but does not force it (most are benign gardens).
+- Mirror ("dextro") = inedible but *guaranteed* inert (can''t infect/poison — safe to study).
+- "Safe" ≠ "edible" — decoupled. Bio-safe is guaranteed on Mirror; food-safe is Compatible-only.
+- Achiral seam (always works either way): breathe air, drink water, mine ore, burn/build with native biomass. Only eat/metabolize/medicate/infect/chirally-poison are gated.
+- ~50/50 Compatible:Mirror is a DESIGN TARGET across canon-silent worlds, not a per-world draw.
+
+WORKSTREAMS
+1. CHIRALITY ASSIGNMENT (Miri). Give every inhabited body a biosphere_class by authored identity: principled default from `economic_base_primary` + `planet_class` (food/drink exporters → Compatible; dead+maintained-atmosphere → TerraformedSterile), explicit overrides where canon speaks. Locked exemplars: Arbour (GJ-338B)=Compatible; Solandar (GJ-827)=Mirror; Keid (GJ-166A)=Compatible (preserved *from exploitation* of edible alien life — the stronger conservation motive; Solandar is the Mirror-inert preserve); Oshima (GJ-79)=TerraformedSterile. Hit the ~50/50 target across the silent remainder.
+2. WIKI CONTINUITY (Miri). Reconcile the entries that already gesture at native biochemistry so each is consistent with its assigned bit + the mechanism: GJ-827, GJ-524, GJ-166A, GJ-684A, GJ-197, GJ-79, GJ-124 (resolve whether Hạ Long''s "protein" is native-Compatible harvest or Earth aquaculture). Additive/explanatory, not contradictory — preserve deliberately-vague lines as intentional.
+3. biosphere_class AUTHORING PATH (asset-pipeline discipline — schema + source + values together, per .claude/rules/asset-pipeline.md). Add `biosphere_class` to wiki body frontmatter (where `substrate:` already lives), a nullable `bodies.biosphere_class` column in systems-schema.sql + MIGRATION_SQL, and importer read. NOT needed for D-246; lands with this pass.
+4. ALIEN-ECOLOGY VOCABULARY (Miri + Araminta) — feeds T-1084 directly. The mosaic palette is keyed on surface-class × climate-biome × biosphere-register (D-246 Palette-key). Provide the micro-habitat NAMES per biome × register: native-mirror niches get *alien* clade names (Solandar''s "five kingdoms" flavor); introduced-Earth niches get Earth names (wheatfield, orchard, pasture). Fill the fixed niche spine (canopy / understory / wet-hollow / pioneer / bare) per climate-biome (savanna gallery-copse, tundra thermokarst / lichen-heath, boreal muskeg / deadfall, …). DETERMINISM: the vocab must be fixed before T-1084 generates its first world (a later addition re-rolls affected voxels). Table size is not a perf/storage concern — author generously but to taste (~100k is headroom, not a target).
+
+BURNELLI (stakeholder, follow-on): a food-supply demand primitive keyed on biosphere_class (Mirror + frontier → import dependency; Compatible → local-food capable). Phase-2 economy consumer; does not block this ticket.
+
+CONSTRAINTS
+- Authored input, not derivation (Arbour''s estate food must never be coin-flipped inedible).
+- Semantic axes stay clade-agnostic (Forest = canopy whether oak or mega-fungus); the register skins niches, it does not change gameplay semantics.
+- The mosaic geometry is chirality-blind — this work does NOT touch voxel geometry, only edibility/economy/naming/vocab.
+
+Chirality decisions resolved (2026-07-02, with Jeroen):
+- Locked exemplars: Arbour=NativeCompatible, Solandar=NativeMirror, Keid=NativeCompatible, Oshima=TerraformedSterile.
+- Miri assignments accepted: GJ-524=Mirror, Freyburg(GJ-684A)=Compatible, Hạ Long(GJ-124)=Compatible (native-Compatible harvest, per her rationale).
+- USER OVERRIDES: Puerto Último(GJ-197)=NativeCompatible (breadbasket identity over Miri''s balance-motivated Mirror). Edict/Sirius(GJ-244A)=Airless — keep the world harsh/frozen (deliberately-inconvenient below-grade transit capital is load-bearing canon); no climate change; below-grade sealed habitat = no open-air biosphere.
+- Two-gate default rule ACCEPTED for canon-silent worlds: Gate 0 (habitability from atmosphere/hydrosphere frontmatter → Airless / TerraformedSterile / Alive), Gate 1 (Compatible vs Mirror from economic_base_primary; agriculture→Compatible near-lock, penal→Mirror, research→50/50 lever).
+- NAMING: Keid''s habitable body renamed Cadwal→(pending final pick; rec Linnaeus — pairs with Solander, its student naturalist; or Beagle). Capital stays ''Vance'' (already the author tribute). Miri''s IP flag (Cadwal = Vance''s exact book title) is the driver.
+- Still to APPLY: the 7 additive wiki continuity edits (Miri specified exact text), the biosphere_class frontmatter+schema+importer plumbing (workstream 3), and the Cadwal→<name> rename across wiki + the GJ166A atlas proposal.
+
+NAMING CONFIRMED: Keid''s habitable body Cadwal → Linnaeus (the great classifier; ties to Solander/Solandar taxonomy + the ''five kingdoms''). Capital stays Vance.
+
+PROGRESS 2026-07-03: Workstreams 1-2 DONE + committed (388168eef, branch micro-habitat-mosaic).
+- Cadwal→Linnaeus rename applied (body frontmatter name + biosphere_class:NativeCompatible, GJ-166A infobox, atlas proposal, D-247 clarified). Capital stays Vance.
+- 5 wiki continuity edits applied (GJ-524 Mirror, GJ-166A/Keid Compatible, GJ-684A Freyburg Compatible, GJ-124 Hạ Long Compatible, GJ-197 Puerto Último Compatible). Solandar/Oshima needed none.
+REMAINING:
+- Workstream 3 (biosphere_class plumbing): add nullable bodies.biosphere_class column to systems-schema.sql; teach tooling/db/wiki_sync.py import_from_wiki to read biosphere_class frontmatter + apply the two-gate DEFAULT for canon-silent bodies (Gate0 atmosphere/hydrosphere, Gate1 economic_role/economic_base); add explicit frontmatter for the 3 override bodies still needing it (Solandar=NativeMirror, GJ-524=NativeMirror [uninhabited-but-alive → default would mis-Airless it], Edict/GJ244Ad=Airless [standard-atm+ice edge case]); ensure generate_wiki round-trips biosphere_class; then the BAKE (run import_from_wiki + generate_wiki → systems.db + regenerated infoboxes, incl. the Linnaeus name propagation) + commit systems.db. Add biosphere_class to BodyParams read set (D-240) as the D-247 code hook. NOTE: the bake is a systems.db binary-artifact commit — consequential, do deliberately.
+- Workstream 4 (alien-ecology vocab → palette): integrate Miri''s biome × register vocabulary (cane/vane/wrack/husk/drum-root native-mirror lexicon + Earth-register names, per her Deliverable 3) into atlas/mosaic.rs + mosaic_constants.toml — extends the temperate/native baseline to the full surface-class × climate-biome × register palette. This also completes T-1084''s biome-awareness. Determinism: lock before generation.
+Miri''s full Deliverable 3 vocab (the source for workstream 4) is in her T-1085 agent report.', NULL, '2026-07-03 20:40:15', '2026-07-03 20:40:15.305', '2026-07-03 20:40:15.305', NULL, 'fa34e073db0be60d59ec01ebd7a5e5f4', 2) ON CONFLICT(hash) DO NOTHING;
diff --git a/.pql/changelog/tickets/2026-07.sql b/.pql/changelog/tickets/2026-07.sql
index 806ece109..4b6af1892 100644
--- a/.pql/changelog/tickets/2026-07.sql
+++ b/.pql/changelog/tickets/2026-07.sql
@@ -218,3 +218,49 @@ Chirality decisions resolved (2026-07-02, with Jeroen):
 - Still to APPLY: the 7 additive wiki continuity edits (Miri specified exact text), the biosphere_class frontmatter+schema+importer plumbing (workstream 3), and the Cadwal→<name> rename across wiki + the GJ166A atlas proposal.
 
 NAMING CONFIRMED: Keid''s habitable body Cadwal → Linnaeus (the great classifier; ties to Solander/Solandar taxonomy + the ''five kingdoms''). Capital stays Vance.', 'in_progress', 'medium', 'miri', NULL, 'D-247', '2026-07-02 20:30:57.222', '2026-07-03 20:20:01.466', NULL, 'f7bd0e6fd21f924f65afcec9a7ed9624', 2) ON CONFLICT(record_id) DO UPDATE SET type=excluded.type, parent_record_id=excluded.parent_record_id, title=excluded.title, description=excluded.description, status=excluded.status, priority=excluded.priority, assigned_to=excluded.assigned_to, team=excluded.team, decision_ref=excluded.decision_ref, updated_at=excluded.updated_at, deleted_at=excluded.deleted_at, hash=excluded.hash, canonical_version=excluded.canonical_version WHERE excluded.updated_at > tickets.updated_at OR (excluded.updated_at = tickets.updated_at AND excluded.hash > tickets.hash);
+INSERT INTO tickets (record_id, type, parent_record_id, title, description, status, priority, assigned_to, team, decision_ref, created_at, updated_at, deleted_at, hash, canonical_version) VALUES ('06FJ91R8RV38BJXGB2V55FP3B8', 'story', '06FGSJNP8YM537DJCC82ZD6544', 'Biosphere register (D-247) — chirality bits, wiki continuity, biosphere_class authoring + alien-ecology vocab', 'Operationalize the D-247 native-biosphere premise. Full brief appended.
+
+---
+BRIEF (2026-07-02) — governing record: D-247 (governance/decisions/content.md); consumer: D-246 / T-1084 mosaic register.
+
+WHAT THIS IS
+D-247 established that habitable Reach worlds were alive before humans (the settlement timeline forbids terraforming from dead rock), and that an AUTHORED per-world chirality bit gates whether native life is edible. This ticket turns that canon into data + reconciled wiki + the vocabulary the mosaic needs. Owner: Miri (worldbuilding/continuity). Stakeholder: Burnelli (food-import economics). Co-maintainer for the vocab: Araminta (visual grammar, per the T-995 ObjectTag precedent).
+
+THE MODEL (locked in D-247 — do not re-litigate, apply it):
+- `biosphere_class ∈ {NativeCompatible, NativeMirror, TerraformedSterile, Airless}`. It is an AUTHORED INPUT, never seed-derived — it follows a world''s economic/cultural identity.
+- Compatible ("levo") = edible; coupling *permits* an authored hazard but does not force it (most are benign gardens).
+- Mirror ("dextro") = inedible but *guaranteed* inert (can''t infect/poison — safe to study).
+- "Safe" ≠ "edible" — decoupled. Bio-safe is guaranteed on Mirror; food-safe is Compatible-only.
+- Achiral seam (always works either way): breathe air, drink water, mine ore, burn/build with native biomass. Only eat/metabolize/medicate/infect/chirally-poison are gated.
+- ~50/50 Compatible:Mirror is a DESIGN TARGET across canon-silent worlds, not a per-world draw.
+
+WORKSTREAMS
+1. CHIRALITY ASSIGNMENT (Miri). Give every inhabited body a biosphere_class by authored identity: principled default from `economic_base_primary` + `planet_class` (food/drink exporters → Compatible; dead+maintained-atmosphere → TerraformedSterile), explicit overrides where canon speaks. Locked exemplars: Arbour (GJ-338B)=Compatible; Solandar (GJ-827)=Mirror; Keid (GJ-166A)=Compatible (preserved *from exploitation* of edible alien life — the stronger conservation motive; Solandar is the Mirror-inert preserve); Oshima (GJ-79)=TerraformedSterile. Hit the ~50/50 target across the silent remainder.
+2. WIKI CONTINUITY (Miri). Reconcile the entries that already gesture at native biochemistry so each is consistent with its assigned bit + the mechanism: GJ-827, GJ-524, GJ-166A, GJ-684A, GJ-197, GJ-79, GJ-124 (resolve whether Hạ Long''s "protein" is native-Compatible harvest or Earth aquaculture). Additive/explanatory, not contradictory — preserve deliberately-vague lines as intentional.
+3. biosphere_class AUTHORING PATH (asset-pipeline discipline — schema + source + values together, per .claude/rules/asset-pipeline.md). Add `biosphere_class` to wiki body frontmatter (where `substrate:` already lives), a nullable `bodies.biosphere_class` column in systems-schema.sql + MIGRATION_SQL, and importer read. NOT needed for D-246; lands with this pass.
+4. ALIEN-ECOLOGY VOCABULARY (Miri + Araminta) — feeds T-1084 directly. The mosaic palette is keyed on surface-class × climate-biome × biosphere-register (D-246 Palette-key). Provide the micro-habitat NAMES per biome × register: native-mirror niches get *alien* clade names (Solandar''s "five kingdoms" flavor); introduced-Earth niches get Earth names (wheatfield, orchard, pasture). Fill the fixed niche spine (canopy / understory / wet-hollow / pioneer / bare) per climate-biome (savanna gallery-copse, tundra thermokarst / lichen-heath, boreal muskeg / deadfall, …). DETERMINISM: the vocab must be fixed before T-1084 generates its first world (a later addition re-rolls affected voxels). Table size is not a perf/storage concern — author generously but to taste (~100k is headroom, not a target).
+
+BURNELLI (stakeholder, follow-on): a food-supply demand primitive keyed on biosphere_class (Mirror + frontier → import dependency; Compatible → local-food capable). Phase-2 economy consumer; does not block this ticket.
+
+CONSTRAINTS
+- Authored input, not derivation (Arbour''s estate food must never be coin-flipped inedible).
+- Semantic axes stay clade-agnostic (Forest = canopy whether oak or mega-fungus); the register skins niches, it does not change gameplay semantics.
+- The mosaic geometry is chirality-blind — this work does NOT touch voxel geometry, only edibility/economy/naming/vocab.
+
+Chirality decisions resolved (2026-07-02, with Jeroen):
+- Locked exemplars: Arbour=NativeCompatible, Solandar=NativeMirror, Keid=NativeCompatible, Oshima=TerraformedSterile.
+- Miri assignments accepted: GJ-524=Mirror, Freyburg(GJ-684A)=Compatible, Hạ Long(GJ-124)=Compatible (native-Compatible harvest, per her rationale).
+- USER OVERRIDES: Puerto Último(GJ-197)=NativeCompatible (breadbasket identity over Miri''s balance-motivated Mirror). Edict/Sirius(GJ-244A)=Airless — keep the world harsh/frozen (deliberately-inconvenient below-grade transit capital is load-bearing canon); no climate change; below-grade sealed habitat = no open-air biosphere.
+- Two-gate default rule ACCEPTED for canon-silent worlds: Gate 0 (habitability from atmosphere/hydrosphere frontmatter → Airless / TerraformedSterile / Alive), Gate 1 (Compatible vs Mirror from economic_base_primary; agriculture→Compatible near-lock, penal→Mirror, research→50/50 lever).
+- NAMING: Keid''s habitable body renamed Cadwal→(pending final pick; rec Linnaeus — pairs with Solander, its student naturalist; or Beagle). Capital stays ''Vance'' (already the author tribute). Miri''s IP flag (Cadwal = Vance''s exact book title) is the driver.
+- Still to APPLY: the 7 additive wiki continuity edits (Miri specified exact text), the biosphere_class frontmatter+schema+importer plumbing (workstream 3), and the Cadwal→<name> rename across wiki + the GJ166A atlas proposal.
+
+NAMING CONFIRMED: Keid''s habitable body Cadwal → Linnaeus (the great classifier; ties to Solander/Solandar taxonomy + the ''five kingdoms''). Capital stays Vance.
+
+PROGRESS 2026-07-03: Workstreams 1-2 DONE + committed (388168eef, branch micro-habitat-mosaic).
+- Cadwal→Linnaeus rename applied (body frontmatter name + biosphere_class:NativeCompatible, GJ-166A infobox, atlas proposal, D-247 clarified). Capital stays Vance.
+- 5 wiki continuity edits applied (GJ-524 Mirror, GJ-166A/Keid Compatible, GJ-684A Freyburg Compatible, GJ-124 Hạ Long Compatible, GJ-197 Puerto Último Compatible). Solandar/Oshima needed none.
+REMAINING:
+- Workstream 3 (biosphere_class plumbing): add nullable bodies.biosphere_class column to systems-schema.sql; teach tooling/db/wiki_sync.py import_from_wiki to read biosphere_class frontmatter + apply the two-gate DEFAULT for canon-silent bodies (Gate0 atmosphere/hydrosphere, Gate1 economic_role/economic_base); add explicit frontmatter for the 3 override bodies still needing it (Solandar=NativeMirror, GJ-524=NativeMirror [uninhabited-but-alive → default would mis-Airless it], Edict/GJ244Ad=Airless [standard-atm+ice edge case]); ensure generate_wiki round-trips biosphere_class; then the BAKE (run import_from_wiki + generate_wiki → systems.db + regenerated infoboxes, incl. the Linnaeus name propagation) + commit systems.db. Add biosphere_class to BodyParams read set (D-240) as the D-247 code hook. NOTE: the bake is a systems.db binary-artifact commit — consequential, do deliberately.
+- Workstream 4 (alien-ecology vocab → palette): integrate Miri''s biome × register vocabulary (cane/vane/wrack/husk/drum-root native-mirror lexicon + Earth-register names, per her Deliverable 3) into atlas/mosaic.rs + mosaic_constants.toml — extends the temperate/native baseline to the full surface-class × climate-biome × register palette. This also completes T-1084''s biome-awareness. Determinism: lock before generation.
+Miri''s full Deliverable 3 vocab (the source for workstream 4) is in her T-1085 agent report.', 'in_progress', 'medium', 'miri', NULL, 'D-247', '2026-07-02 20:30:57.222', '2026-07-03 20:40:15.305', NULL, '83246ebc2aa0550f44afd68721efb9d4', 2) ON CONFLICT(record_id) DO UPDATE SET type=excluded.type, parent_record_id=excluded.parent_record_id, title=excluded.title, description=excluded.description, status=excluded.status, priority=excluded.priority, assigned_to=excluded.assigned_to, team=excluded.team, decision_ref=excluded.decision_ref, updated_at=excluded.updated_at, deleted_at=excluded.deleted_at, hash=excluded.hash, canonical_version=excluded.canonical_version WHERE excluded.updated_at > tickets.updated_at OR (excluded.updated_at = tickets.updated_at AND excluded.hash > tickets.hash);
diff --git a/docs/design/alien-ecology-vocabulary.md b/docs/design/alien-ecology-vocabulary.md
new file mode 100644
index 000000000..dffcc10f4
--- /dev/null
+++ b/docs/design/alien-ecology-vocabulary.md
@@ -0,0 +1,202 @@
+# Alien-Ecology Vocabulary — micro-habitat naming layer (D-247 / D-246 / T-1085)
+
+Authored by **Miri** (T-1085 workstream 4), 2026-07-03. This is the **naming /
+rendering** layer that skins the mosaic's semantic micro-habitats (T-1084, D-246).
+The mosaic selects **D-228 axis-triples** (terrain × vegetation × water); this table
+supplies the **display names** per `climate-biome × register × niche`, so a "canopy"
+niche in a temperate forest reads as *vane-cane stand* (native-mirror) or *dense
+stand* (introduced-Earth). Names are **derived labels** (D-228) — never stored on the
+voxel. Co-maintenance with **Araminta** for visual grammar (per the T-995 ObjectTag
+precedent).
+
+Governing canon: [D-247](../../governance/decisions/content.md) (native biospheres +
+chirality), [D-246](../../governance/decisions/architecture.md) (the mosaic that
+consumes this). Determinism: this vocabulary is part of the deterministic input
+surface once wired — fix it before the mosaic generates its first world.
+
+---
+
+## Methodology
+
+### Niche spine (the cross-biome authoring scaffold — reinterpreted per biome)
+
+The five niches are **roles**, not literal "trees exist here" claims — each biome
+reinterprets them (a grassland's "canopy" is its tallest grass stratum):
+
+- **canopy** — the biome's tallest/densest cover stratum. Full overhead concealment
+  where the biome has one; the thickest topmost growth otherwise. Blocks sightlines +
+  weather; sets the movement-sound baseline.
+- **understory** — the layer beneath/within canopy, or a secondary partial-cover
+  stratum where there's no true canopy. Partial concealment, filtered light.
+- **wet-hollow** — moisture-conditioned low point, tied to the `voxel_relief`
+  micro-thalweg (D-246). Slows movement; sound and smell carry differently.
+- **pioneer** — colonizing / disturbed / regrowth ground; first-succession or edge
+  margin. The "in-between" transition zone.
+- **bare-ground** — minimal-to-no cover, exposed substrate. Best sightlines, least
+  concealment, fastest movement; the biome's "clearing"/"pan" default.
+
+### Register naming
+
+- **Introduced-Earth** — ordinary recognizable agricultural/ecological English
+  (orchard, pasture, hedgerow): warm, human-scaled, legible at a glance.
+- **Native (alien)** — built from a small recurring invented growth-habit lexicon (the
+  alien equivalent of "tree / grass / moss / reed" as informal field terms, not formal
+  taxonomy — Solandar's "five kingdoms" stays deliberately unnamed at the formal level).
+
+### The native growth-habit lexicon
+
+| Root | Fills the role of | Typical niche |
+|---|---|---|
+| **cane** | tall, upright, single-or-clustered stalks | canopy / understory in wooded biomes |
+| **vane** | broad, blade-like, often iridescent structures | canopy (the richest biomes) |
+| **wrack** | low, spreading, mat-forming growth | bare-ground / pioneer / understory |
+| **husk** | bulb/pod-bearing, clustered near water | wet-hollow |
+| **drum-root** | colonial, ring-spreading root-network | pioneer (recurs across Grassland/Tundra/Savanna) |
+
+A few names deliberately **recur across biomes** (husk-fall tangle, lichen-scald flat,
+scorch-bloom flush, the drum-root/ring motif) — intentional coherence: it reads as one
+designed ecology, not eleven unrelated ones. This is the Bobiverse "convergent-but-alien
+niches" touchstone made concrete: the same growth-habit fills the same niche across
+biomes, as oak/pine/baobab all fill "canopy" on Earth without being the same species.
+
+### Sensory discipline — the achiral seam (D-247) stays intact
+
+Native water is always safe to drink, ground always safe to mine/walk, biomass always
+safe to burn/build. Every note flavors **strangeness** (smell, sound, texture, footing),
+never *danger*, for those actions. Where hazard flavor appears it is **mechanical/
+textural** (brittle husks giving way, thorns, uneven footing) — never biochemical
+(Mirror life can't infect or poison; Compatible hazard is an *authored* option, flagged
+as a hook, not exercised on any specific world here).
+
+### Scope
+
+This table fills **climate-biome × register**, richest on the six vegetated biomes,
+lighter on the five extreme ones (niches merged where they don't naturally apply). A
+full `climate-biome × surface-class × register` cross-product is a future pass; the
+D-246 Sand/Rock/Gravel/Lava surface-class rows already carry register-neutral flavor.
+
+---
+
+## Rich biomes
+
+### TropicalWet
+*The Solandar register — hot, wet, maximum biomass, fast decomposition. "Twelve thousand species, five kingdoms."*
+
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Canopy | **Vane-crown stand** — towering oil-sheened vane-blades interlock into a solid ceiling; understory twilight-dim at midday. A thick sweetish rot-and-sap smell that never clears. | **Cacao shade-block** — planted tree-crop under managed shade canopy; rows just regular enough to give away human hands. |
+| Understory | **Wrack-drape thicket** — wet curtains of wrack draping from the canopy; clings and slows movement, leaves a faint chemical tang. | **Shade-coffee row** — broad-leafed cultivars beneath the plantation canopy. |
+| Wet-hollow | **Husk-seep hollow** — bellied husk-growths at a warm standing seep; water runs faintly opalescent with alien tannins — safe to drink, unmistakably not Earth's water by the smell. | **Rice-paddy terrace** — flooded cultivated wet-ground rows. |
+| Pioneer | **Scorch-bloom flush** — fast alien color-succession flooding a canopy-fall light-gap within weeks, before slower vane-forms retake it. | **Cassava clearing** — a slashed-and-fallow plot gone to quick Earth pioneer weeds and root crops. |
+| Bare-ground | **Salt-crust flat** — mineral-crusted open patch where nothing native roots, by chemistry not disturbance; wrong-colored crust crunches underfoot. | **Threshing yard** — bare packed earth cut into the plantation. |
+
+### TemperateForest
+
+| Niche | Native-Mirror | Introduced-Earth (existing D-246 baseline) |
+|---|---|---|
+| Canopy | **Vane-cane stand** — cane trunks crowned with stiff, faintly metallic-blue vane-fronds; a dry clicking rustle, not a hiss — it doesn't *sound* like a forest. | **Dense stand** |
+| Understory | **Wrack-frond undergrowth** — knee-high tangle, cool and slightly rubbery; brushing it releases a sharp resinous smell with no Earth analog. | **Fern undergrowth** |
+| Wet-hollow | **Husk-reed run** — husk-reeds trace the brook's micro-thalweg; husks pop and refill audibly as current passes — heard before seen. | **Brook** |
+| Pioneer | **Husk-fall tangle** — collapsed cane-husks, brittle and hollow rather than woody; lighter to move than deadfall but sharper underfoot (a footing hazard, not a fire-fuel one). | **Deadfall** |
+| Bare-ground | **Bloom-scald clearing** — kept open by an allelopathic wrack that poisons competing roots (achiral-safe to walk, just barren); faint bitter-almond air. | **Glade** |
+
+### TemperateGrassland
+
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Canopy | **Cane-tassel sweep** — waist-to-shoulder cane tipped with heavy seed-tassels; moves in slow grass-like waves, but the tassels chime softly against each other. | **Tussock** |
+| Understory | **Wrack-bloom understory** — a low flowering layer threading between the cane-tassels; a sweet "almost right" smell that unsettles Earth-register visitors. | **Wildflower** |
+| Wet-hollow | **Husk-line seep** — the ephemeral channel lined with squat husk-forms that only surface after rain — a visible tell of where the water runs even in dry season. | **Dry wash** |
+| Pioneer | **Drum-root knot** — a knotted ring of colonial root-growth pushing up through the grass, spreading centimeters a year; the ring's center is always slightly bare. | **Scrub island** |
+| Bare-ground | **Hardpan scald** — a mineral-crusted bare patch; the ground rings slightly hollow underfoot. | **Grazing pan** — bare-trodden earth from stock. |
+
+### BorealForest
+
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Canopy | **Rime-vane stand** — slow, frost-hardy vane-cane holding a permanent rime-like bloom even in summer; dim, cold, nearly silent — a "wrong" quiet. | **Spruce-shadow stand** — conifer-analog, deep cold shade, needle-duff footing. |
+| Understory | **Wrack-moss carpet** — thick spongy wrack holding meltwater like a sponge; boots sink and release a cold mineral smell. | **Bilberry scrub** |
+| Wet-hollow | **Husk-bog muskeg** — waterlogged cold ground where the tussocks are husk-forms; each step risks a hollow husk giving way. | **Muskeg fen** |
+| Pioneer | **Husk-fall tangle** (shared with TemperateForest) | **Deadfall** — cold-climate blowdown. |
+| Bare-ground | **Lichen-scald flat** — thin pale native crust over permafrost-like ground, ancient and slow; disturbing it outlasts a human lifetime. | **Burn scar** |
+
+### Tundra
+*Treeless, thermokarst. Thin biomass; niches all present but small.*
+
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Canopy (dwarf) | **Husk-heath brake** — the tallest native growth is ankle-high, a dense husk-mat; reads as heath at distance, clearly not-leaves up close. | **Dwarf-willow brake** |
+| Understory | **Wrack-lichen mat** — a thin crust; grows so slowly that a season's footpath shows a decade later. | **Moss-sedge mat** |
+| Wet-hollow | **Thermokarst husk-pool** — a collapse-and-thaw pool ringed with husk-forms that emerge only in the brief thaw window — seasonal in a way no other biome's wet-hollow is. | **Thaw-pond sedge** |
+| Pioneer | **Drum-root ring** (shared motif, at tundra scale — centimeters across after decades) | **Gravel scar** |
+| Bare-ground | **Lichen-scald flat** (shared with Boreal — here over patterned-ground polygons) | **Frost-heave flat** |
+
+### Savanna
+*Warm, seasonal, grass-dominant with scattered tree-copses.*
+
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Canopy | **Vane-crown copse** — an isolated stand, the only real shade for kilometers; mistakable for a gallery copse at distance, but the bark-analog is smooth and faintly warm. | **Gallery copse** |
+| Understory | **Husk-thorn scrub** — thorny native scrub, identical obstruction/concealment to acacia; the thorns are hollow and whistle faintly in wind. | **Acacia scrub** |
+| Wet-hollow | **Husk-pan waterhole** — seasonal flood-and-dry; dormant husk-forms visibly "wake" within days of the first rain. | **Seasonal waterhole** |
+| Pioneer | **Scorch-bloom flush** (shared with TropicalWet — succession by seasonal fire) | **Burnt-grass flush** |
+| Bare-ground | **Drum-mound flat** — a hardpan clearing built by colonial drum-root activity; bare, sun-baked, faintly warm underfoot even at night. | **Termite-pan clearing** |
+
+---
+
+## Lighter biomes (extreme classes — niches merged/noted where they don't apply)
+
+### Desert
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Canopy ("tallest thing here") | **Cane-spire stand** — sparse deep-taprooted spires, each commanding the horizon with a thin moving shadow. | **Date-palm draw** |
+| Understory + pioneer (merged) | **Wrack-thorn scatter** — sparse widely-spaced domed clumps hoarding moisture. | **Scrub flat** |
+| Wet-hollow | **Husk-spring hollow** — a rare seep ringed with water-swollen husk-forms; the local landmark. Water fine to drink; nothing in the ring looks edible. | **Wadi oasis** |
+| Bare-ground | *served by the D-246 Sand/Rock rows* (dune crest, hardpan, outcrop, scree) — bare rock/sand carries no register. | (same) |
+
+### Alpine
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Canopy (treeline fringe) | **Wind-cane fringe** — wind-flagged, permanently bent one way; the flag-shape wayfinds like krummholz, the material reads wrong up close. | **Krummholz fringe** |
+| Understory + pioneer (merged) | **Wrack-cushion field** — tight ground-hugging growth, faintly warm to bare skin (a metabolic-heat tell Earth cushion plants lack). | **Cushion-plant field** |
+| Wet-hollow | **Husk-melt seep** — seasonal meltwater seep, husk-forms present only weeks a year. | **Snowmelt sedge** |
+| Bare-ground | *served by the Rock rows* (outcrop/scree/talus/ledge/**lichen** — lichen doubles as this biome's dominant cover). | (same) |
+
+### Volcanic
+*D-210 flags this the least data-grounded biome; the Lava surface-class already has the richest table. Kept light.*
+
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Recovering slope (canopy/understory collapsed) | **Husk-root slope** — pioneer husk-root colonizing cooled ash, first native life back after a flow. | **Ash-orchard slope** — Earth fruit trees on fertile volcanic ash. |
+| Wet-hollow | **Husk-vent seep** — a geothermal seep, the hardiest husk-forms in the register; warm, safe to bathe in. | **Hot-spring terrace** |
+| Pioneer + bare-ground | *served by the Lava rows* (fresh sheet, weathered crust w/ Lichen, tube collapse, pioneer scrub). | (same) |
+
+### Wetland (climate-biome)
+*Heavily overlaps the surface-class Wetland table (richest baseline). Native mirror given for symmetry; kept light.*
+
+| Niche | Native-Mirror | Introduced-Earth (existing D-246 names) |
+|---|---|---|
+| Canopy | **Husk-crown copse** — water-rooted husk-trees hung with drip-tendrils. | **Copse** |
+| Understory / bare-ground | **Wrack-flat clearing** — open wrack-carpeted shallows. | **Meadow / Clearing** |
+| Wet-hollow (×3) | **Husk-bog / Husk-marsh / Husk-run** — the marsh variant's reed-analog hisses faintly when disturbed, a natural detection tell. | **Bog / Marsh / Creek** |
+| Pioneer | thin/underdeveloped (same gap as the baseline) | (same gap) |
+
+### CoastalLowland
+| Niche | Native-Mirror | Introduced-Earth |
+|---|---|---|
+| Canopy | **Vane-palm fringe** — salt-tolerant, leaning permanently seaward; same visual tell as a coastal treeline, different material. | **Coconut fringe** |
+| Understory + pioneer (merged) | **Wrack-tangle scrub** — salt-hardy tangle; smells of ozone and brine-that's-not-quite-brine. | **Dune scrub** |
+| Wet-hollow (tidal) | **Husk-tide flat** — husk-forms seal shut at low tide and visibly "breathe" open as water returns — a legible tide-state tell. | **Salt-marsh flat** |
+| Bare-ground | **Husk-shard beach** — the tideline wrack is broken husk-fragments instead of shells; the wrong shapes up close. | **Shell-sand beach** |
+
+---
+
+## Open hooks (Miri, T-1085)
+
+- **Five kingdoms** — Solandar's "twelve thousand species across five kingdoms" is left
+  formally unnamed (deliberately-vague canon). Naming the kingdoms is a future hook.
+- **Compatible-world hazard** — a Compatible (levo) biome (TropicalWet, Savanna) is the
+  right *register* for a future authored native pathogen/toxin (D-247 §"coupling permits
+  but doesn't force"); a hook, not yet exercised.
+- **Full cross-product** — climate-biome × **surface-class** × register (this table is
+  climate-biome × register only) is a future pass.
diff --git a/server/data/mosaic_constants.toml b/server/data/mosaic_constants.toml
index 0bd689dc5..7228d2feb 100644
--- a/server/data/mosaic_constants.toml
+++ b/server/data/mosaic_constants.toml
@@ -107,38 +107,160 @@ water = "Dry"
 weight = 15
 affinity = "any"
 
-[[grassland]]
-name = "tussock"
+# Grassland is BIOME-KEYED (T-1085, D-246): the ClimateBiome derived from the district's
+# temperature_c + moisture_q selects savanna / temperate / boreal / tundra, so a
+# grassland reads distinctly per climate rather than as one biome-blind palette. §8-safe
+# (vegetation + water; terrain stays Soil in v1). Names per
+# docs/design/alien-ecology-vocabulary.md.
+
+[[grassland_temperate]]
+name = "cane-tassel sweep"
 terrain = "Soil"
 vegetation = "Grass"
 water = "Dry"
 weight = 40
 affinity = "any"
 
-[[grassland]]
-name = "scrub island"
+[[grassland_temperate]]
+name = "drum-root knot"
 terrain = "Soil"
 vegetation = "Scrub"
 water = "Dry"
 weight = 18
 affinity = "dry"
 
-[[grassland]]
-name = "wildflower"
+[[grassland_temperate]]
+name = "wrack-bloom"
 terrain = "Soil"
 vegetation = "Meadow"
 water = "Dry"
 weight = 17
 affinity = "dry"
 
-[[grassland]]
-name = "dry wash"
-terrain = "Gravel"
+[[grassland_temperate]]
+name = "husk-line seep"
+terrain = "Soil"
 vegetation = "Barren"
 water = "Dry"
 weight = 10
 affinity = "wet"
 
+# Savanna: scattered Forest copses in a grass matrix — the vane-crown copse is the
+# clearest distinction from temperate grassland.
+[[grassland_savanna]]
+name = "cane-tassel sweep"
+terrain = "Soil"
+vegetation = "Grass"
+water = "Dry"
+weight = 38
+affinity = "any"
+
+[[grassland_savanna]]
+name = "vane-crown copse"
+terrain = "Soil"
+vegetation = "Forest"
+water = "Dry"
+weight = 12
+affinity = "dry"
+
+[[grassland_savanna]]
+name = "husk-thorn scrub"
+terrain = "Soil"
+vegetation = "Scrub"
+water = "Dry"
+weight = 18
+affinity = "dry"
+
+[[grassland_savanna]]
+name = "husk-pan waterhole"
+terrain = "Soil"
+vegetation = "Grass"
+water = "Shallow"
+weight = 10
+affinity = "wet"
+
+[[grassland_savanna]]
+name = "drum-mound flat"
+terrain = "Soil"
+vegetation = "Barren"
+water = "Dry"
+weight = 12
+affinity = "dry"
+
+# Boreal grassland: cooler, sparser — more scrub, a lichen fraction, no meadow.
+[[grassland_boreal]]
+name = "cane-tassel sweep"
+terrain = "Soil"
+vegetation = "Grass"
+water = "Dry"
+weight = 30
+affinity = "any"
+
+[[grassland_boreal]]
+name = "wrack-scrub"
+terrain = "Soil"
+vegetation = "Scrub"
+water = "Dry"
+weight = 25
+affinity = "dry"
+
+[[grassland_boreal]]
+name = "husk-line seep"
+terrain = "Soil"
+vegetation = "Grass"
+water = "Shallow"
+weight = 10
+affinity = "wet"
+
+[[grassland_boreal]]
+name = "wrack-lichen mat"
+terrain = "Soil"
+vegetation = "Lichen"
+water = "Dry"
+weight = 12
+affinity = "any"
+
+[[grassland_boreal]]
+name = "cold scald"
+terrain = "Soil"
+vegetation = "Barren"
+water = "Dry"
+weight = 15
+affinity = "dry"
+
+# Tundra: sparse and cold — scrub, lichen, bare patterned ground, NO forest.
+[[grassland_tundra]]
+name = "husk-heath brake"
+terrain = "Soil"
+vegetation = "Scrub"
+water = "Dry"
+weight = 26
+affinity = "any"
+
+[[grassland_tundra]]
+name = "wrack-lichen mat"
+terrain = "Soil"
+vegetation = "Lichen"
+water = "Dry"
+weight = 24
+affinity = "any"
+
+[[grassland_tundra]]
+name = "thermokarst husk-pool"
+terrain = "Soil"
+vegetation = "Grass"
+water = "Shallow"
+weight = 10
+affinity = "wet"
+
+[[grassland_tundra]]
+name = "lichen-scald flat"
+terrain = "Soil"
+vegetation = "Barren"
+water = "Dry"
+weight = 30
+affinity = "dry"
+
 # ── Follow-up (§8-aware material pass): Sand / Rock / Gravel / Lava ──────────────
 
 [[sand]]
diff --git a/server/src/atlas/mosaic.rs b/server/src/atlas/mosaic.rs
index da8a90fbc..e2414f1b4 100644
--- a/server/src/atlas/mosaic.rs
+++ b/server/src/atlas/mosaic.rs
@@ -85,6 +85,40 @@ impl SurfaceClass {
     }
 }
 
+/// The district's climate-biome (T-1085, D-246) — derived from `temperature_c` +
+/// `moisture_q`. It refines the *vegetated* surface-class palettes so a Grassland reads
+/// as savanna / temperate / boreal / tundra rather than one biome-blind palette (the
+/// D-246 biome-awareness fix). The alien display names per biome × register live in
+/// `docs/design/alien-ecology-vocabulary.md`; this layer carries only the D-228 axes.
+#[derive(Debug, Clone, Copy, PartialEq, Eq)]
+pub enum ClimateBiome {
+    Tropical,
+    Temperate,
+    Boreal,
+    Tundra,
+    Savanna,
+}
+
+impl ClimateBiome {
+    /// Coarse biome from mean-annual temperature (°C) + district moisture (0–100).
+    /// Airless districts never reach the mosaic; the `10.0` default is a safe
+    /// temperate fallback for a missing temperature.
+    pub fn from_climate(temperature_c: Option<f32>, moisture_q: i32) -> Self {
+        let t = temperature_c.unwrap_or(10.0);
+        if t < 0.0 {
+            ClimateBiome::Tundra
+        } else if t < 7.0 {
+            ClimateBiome::Boreal
+        } else if t <= 20.0 {
+            ClimateBiome::Temperate
+        } else if moisture_q >= 45 {
+            ClimateBiome::Tropical
+        } else {
+            ClimateBiome::Savanna
+        }
+    }
+}
+
 // Shorthand for the const tables below.
 use MoistureAffinity::{Any, Dry, Wet};
 use TerrainMaterial as T;
@@ -128,11 +162,49 @@ const FOREST: &[MicroHabitat] = &[
     h("fern undergrowth", T::Soil, V::Thicket, W::Dry, 15, Any),
 ];
 
-const GRASSLAND: &[MicroHabitat] = &[
-    h("tussock", T::Soil, V::Grass, W::Dry, 40, Any),
-    h("scrub island", T::Soil, V::Scrub, W::Dry, 18, Dry),
-    h("wildflower", T::Soil, V::Meadow, W::Dry, 17, Dry),
-    h("dry wash", T::Gravel, V::Barren, W::Dry, 10, Wet),
+// Grassland is biome-keyed (T-1085): savanna / temperate / boreal / tundra read
+// distinctly rather than as one biome-blind palette (D-246). §8-safe — vegetation +
+// water only; terrain stays Soil in v1 (the Gravel dry-wash material variant is the
+// §8-aware follow-up). Names per docs/design/alien-ecology-vocabulary.md.
+const GRASSLAND_TEMPERATE: &[MicroHabitat] = &[
+    h("cane-tassel sweep", T::Soil, V::Grass, W::Dry, 40, Any),
+    h("drum-root knot", T::Soil, V::Scrub, W::Dry, 18, Dry),
+    h("wrack-bloom", T::Soil, V::Meadow, W::Dry, 17, Dry),
+    h("husk-line seep", T::Soil, V::Barren, W::Dry, 10, Wet),
+];
+
+// Savanna: the signature is scattered Forest copses in a grass matrix (the
+// vane-crown copse) — the clearest biome distinction from temperate grassland.
+const GRASSLAND_SAVANNA: &[MicroHabitat] = &[
+    h("cane-tassel sweep", T::Soil, V::Grass, W::Dry, 38, Any),
+    h("vane-crown copse", T::Soil, V::Forest, W::Dry, 12, Dry),
+    h("husk-thorn scrub", T::Soil, V::Scrub, W::Dry, 18, Dry),
+    h("husk-pan waterhole", T::Soil, V::Grass, W::Shallow, 10, Wet),
+    h("drum-mound flat", T::Soil, V::Barren, W::Dry, 12, Dry),
+];
+
+// Boreal grassland: cooler and sparser — more scrub, a lichen fraction, no meadow.
+const GRASSLAND_BOREAL: &[MicroHabitat] = &[
+    h("cane-tassel sweep", T::Soil, V::Grass, W::Dry, 30, Any),
+    h("wrack-scrub", T::Soil, V::Scrub, W::Dry, 25, Dry),
+    h("husk-line seep", T::Soil, V::Grass, W::Shallow, 10, Wet),
+    h("wrack-lichen mat", T::Soil, V::Lichen, W::Dry, 12, Any),
+    h("cold scald", T::Soil, V::Barren, W::Dry, 15, Dry),
+];
+
+// Tundra: sparse and cold — scrub, lichen, bare patterned ground, NO forest.
+const GRASSLAND_TUNDRA: &[MicroHabitat] = &[
+    h("husk-heath brake", T::Soil, V::Scrub, W::Dry, 26, Any),
+    h("wrack-lichen mat", T::Soil, V::Lichen, W::Dry, 24, Any),
+    h(
+        "thermokarst husk-pool",
+        T::Soil,
+        V::Grass,
+        W::Shallow,
+        10,
+        Wet,
+    ),
+    h("lichen-scald flat", T::Soil, V::Barren, W::Dry, 30, Dry),
 ];
 
 const SAND: &[MicroHabitat] = &[
@@ -161,12 +233,19 @@ const LAVA: &[MicroHabitat] = &[
     h("pioneer scrub", T::Lava, V::Scrub, W::Dry, 20, Dry),
 ];
 
-/// The embedded palette for a surface-class (temperate / native baseline).
-pub fn palette(class: SurfaceClass) -> &'static [MicroHabitat] {
+/// The embedded native palette for a `(surface-class, climate-biome)` (D-246 Palette
+/// key). Grassland is biome-keyed (T-1085); the other classes are biome-invariant in
+/// v1 — Forest biomes (tropical/boreal) and the extreme classes are the follow-up.
+pub fn palette(class: SurfaceClass, biome: ClimateBiome) -> &'static [MicroHabitat] {
     match class {
         SurfaceClass::Wetland => WETLAND,
         SurfaceClass::Forest => FOREST,
-        SurfaceClass::Grassland => GRASSLAND,
+        SurfaceClass::Grassland => match biome {
+            ClimateBiome::Savanna | ClimateBiome::Tropical => GRASSLAND_SAVANNA,
+            ClimateBiome::Boreal => GRASSLAND_BOREAL,
+            ClimateBiome::Tundra => GRASSLAND_TUNDRA,
+            ClimateBiome::Temperate => GRASSLAND_TEMPERATE,
+        },
         SurfaceClass::Sand => SAND,
         SurfaceClass::Rock => ROCK,
         SurfaceClass::Gravel => GRAVEL,
@@ -240,7 +319,7 @@ mod tests {
 
     #[test]
     fn every_palette_nonempty_and_positively_weighted() {
-        for class in [
+        let classes = [
             SurfaceClass::Wetland,
             SurfaceClass::Forest,
             SurfaceClass::Grassland,
@@ -248,19 +327,58 @@ mod tests {
             SurfaceClass::Rock,
             SurfaceClass::Gravel,
             SurfaceClass::Lava,
-        ] {
-            let p = palette(class);
-            assert!(!p.is_empty(), "{class:?} palette empty");
-            assert!(
-                p.iter().all(|hab| hab.weight > 0),
-                "{class:?} has a non-positive weight"
-            );
+        ];
+        let biomes = [
+            ClimateBiome::Tropical,
+            ClimateBiome::Temperate,
+            ClimateBiome::Boreal,
+            ClimateBiome::Tundra,
+            ClimateBiome::Savanna,
+        ];
+        for class in classes {
+            for biome in biomes {
+                let p = palette(class, biome);
+                assert!(!p.is_empty(), "{class:?}/{biome:?} palette empty");
+                assert!(
+                    p.iter().all(|hab| hab.weight > 0),
+                    "{class:?}/{biome:?} has a non-positive weight"
+                );
+            }
         }
     }
 
+    #[test]
+    fn from_climate_bands() {
+        use ClimateBiome::*;
+        assert_eq!(ClimateBiome::from_climate(Some(-10.0), 40), Tundra);
+        assert_eq!(ClimateBiome::from_climate(Some(3.0), 40), Boreal);
+        assert_eq!(ClimateBiome::from_climate(Some(12.0), 40), Temperate);
+        assert_eq!(ClimateBiome::from_climate(Some(28.0), 70), Tropical);
+        assert_eq!(ClimateBiome::from_climate(Some(28.0), 20), Savanna);
+    }
+
+    #[test]
+    fn grassland_biomes_are_distinct() {
+        // Savanna carries scattered Forest copses; tundra carries none but has Lichen.
+        let savanna = palette(SurfaceClass::Grassland, ClimateBiome::Savanna);
+        let tundra = palette(SurfaceClass::Grassland, ClimateBiome::Tundra);
+        assert!(
+            savanna.iter().any(|h| h.vegetation == V::Forest),
+            "savanna should have Forest copses"
+        );
+        assert!(
+            !tundra.iter().any(|h| h.vegetation == V::Forest),
+            "tundra must not carry Forest"
+        );
+        assert!(
+            tundra.iter().any(|h| h.vegetation == V::Lichen),
+            "tundra should have Lichen"
+        );
+    }
+
     #[test]
     fn select_is_deterministic_and_spans_the_palette() {
-        let p = palette(SurfaceClass::Wetland);
+        let p = palette(SurfaceClass::Wetland, ClimateBiome::Temperate);
         assert_eq!(select(p, 0.37, 0.5).name, select(p, 0.37, 0.5).name);
         let names: std::collections::BTreeSet<_> = (0..100)
             .map(|i| select(p, i as f64 / 100.0, 0.5).name)
@@ -274,7 +392,7 @@ mod tests {
 
     #[test]
     fn wetness_favours_wet_habitats_over_dry_signal() {
-        let p = palette(SurfaceClass::Wetland);
+        let p = palette(SurfaceClass::Wetland, ClimateBiome::Temperate);
         let wet_count = |wetness: f64| {
             (0..100)
                 .filter(|i| {
diff --git a/server/src/atlas/voxel.rs b/server/src/atlas/voxel.rs
index 46311db1d..b561095a5 100644
--- a/server/src/atlas/voxel.rs
+++ b/server/src/atlas/voxel.rs
@@ -530,10 +530,22 @@ pub fn derive_voxel_column(
         use crate::atlas::mosaic::{self, SurfaceClass};
         let dominant_veg = Vegetation::from_vegetation_class(district_eff.vegetation_class);
         let class = SurfaceClass::from_base(column.terrain, dominant_veg);
+        // Climate-biome (T-1085) refines the Grassland palette: savanna / temperate /
+        // boreal / tundra read distinctly (D-246 biome-awareness).
+        let biome =
+            mosaic::ClimateBiome::from_climate(district_eff.temperature_c, district_eff.moisture_q);
         let apply = match class {
             SurfaceClass::Wetland | SurfaceClass::Forest => true,
-            // Barren grassland (cold / arid) stays barren — no imposed grass.
-            SurfaceClass::Grassland => !matches!(dominant_veg, Vegetation::Barren),
+            // A vegetated grassland always; a barren-dominant one only when the biome is
+            // cold (tundra / boreal), where the sparse lichen/scrub mosaic is climate-
+            // appropriate — a hot barren district (desert on Soil) stays bare.
+            SurfaceClass::Grassland => {
+                !matches!(dominant_veg, Vegetation::Barren)
+                    || matches!(
+                        biome,
+                        mosaic::ClimateBiome::Tundra | mosaic::ClimateBiome::Boreal
+                    )
+            }
             // Rock / Sand / Gravel / Lava are material-driven — a §8-aware follow-up.
             _ => false,
         };
@@ -550,7 +562,7 @@ pub fn derive_voxel_column(
             // lowered on the rises (relief_signal negative in valleys).
             let moisture = (district_eff.moisture_q as f64 / 100.0).clamp(0.0, 1.0);
             let wetness = (moisture - relief_signal).clamp(0.0, 1.0);
-            let hab = mosaic::select(mosaic::palette(class), field, wetness);
+            let hab = mosaic::select(mosaic::palette(class, biome), field, wetness);
             column.vegetation = hab.vegetation;
             // Add standing water only where the family left it dry AND the chunk's
             // hydrology gate allows it (T-1040: water placement must agree with
diff --git a/server/tests/golden/believability.json b/server/tests/golden/believability.json
index da2796ca3..380899855 100644
--- a/server/tests/golden/believability.json
+++ b/server/tests/golden/believability.json
@@ -37,7 +37,7 @@
       "drainage_samples": 0,
       "drainage_monotonic": 0,
       "vegetation_samples": 64,
-      "vegetated_districts": 8
+      "vegetated_districts": 20
     }
   },
   {
@@ -70,7 +70,7 @@
       "vegetation_classes": 1,
       "terrain_materials": 4,
       "voxel_relief_m": 19,
-      "micro_habitat_distinct": 0
+      "micro_habitat_distinct": 1
     },
     "coherence": {
       "water_districts": 25,
@@ -78,7 +78,7 @@
       "drainage_samples": 0,
       "drainage_monotonic": 0,
       "vegetation_samples": 64,
-      "vegetated_districts": 0
+      "vegetated_districts": 39
     }
   }
 ]